why do carnivores have high assimilation efficiency

why do carnivores have high assimilation efficiency

https://doi.org/10.1007/978-1-4419-9504-9_10, DOI: https://doi.org/10.1007/978-1-4419-9504-9_10, eBook Packages: Biomedical and Life SciencesBiomedical and Life Sciences (R0). They are obtained from the diet and are necessary for the proper growth and development of the body. Ecology of a grazing ecosystem: The Serengeti. Bryant, J.P. and P.J. Wiley is a global provider of content and content-enabled workflow solutions in areas of scientific, technical, medical, and scholarly research; professional development; and education. How/why would assimilation efficiency differ for herbivores eating grass, herbivores eating seeds, and carnivores eating herbivores? Barboza, P.S., K.L. Repeated in words, CE is the percentage of total productivity available at one trophic level (Pn-1) that is actually consumed ('ingested') by a trophic compartment 'one level up' (In). Do carnivores have higher assimilation efficiency? 1999. Entrepreneurship Hustle Real Success Course, Energy flow through contrasting communities. However, only about 10% of the total energy from the plant gets stored in the herbivores body as extra body tissue. Example scenario: A farmer wants to sell part of his land. Community structure, population control and competition. Both herbivores and carnivores evolved over millions of years to do two things in their own specific way: survive (and hopefully) reproduce. Assimilation efficiency is the amount of assimilation divided by the amount of food ingestion ( ) Net Production efficiency is the amount of consumer production divided by the amount of assimilation ( ). Is there a database for insurance claims? Frank, D.A. Elliott, J.C. Moore, et al. Food Webs. To complete this calculation, we divide the amount from the higher trophic level by the amount from the lower trophic level and multiply by one hundred. Piaget believed that cognitive development occurs as a result of two processes: adaptation and equilibrium. What is a trophic cascade? This cookie is set by GDPR Cookie Consent plugin. The individual that will pay the most for the land uses the resource more efficiently than someone who does not pay the most money for the land. Pace, M.L., J.J. Cole, S.R. It does not store any personal data. McKey, D., P.G. Berenbaum. Assimilation Efficiency 3.) 1992. Complex trophic interactions in deserts: An empirical critique of food-web theory. Beckerman. Perspectives on Plant Competition. Kitchell. A hypercarnivore is an organism that depends on animals for at least 70 percent of its diet. Ecological efficiency describes the efficiency with which energy is transferred from one trophic level to the next. What are the five methods of dispute resolution? Production Efficiency = (13.3 / 16.7) x 100 = 79.6%, Energy & nutrient transfers between trophic levels. Why might assimilation efficiencies be much higher for herbivores eating seeds than for herbivores eating leaves? Resource availability and plant anti-herbivore defense. Grazing as an optimization process: Grass-ungulate relationships in the Serengeti. Klein. On average, invertebrates use only ~80% of assimilated energy for metabolism, and thus exhibit greater net production efficiency (~20%) than do vertebrates. How do you calculate energy efficiency in a food chain? She or he will best know the preferred format. In ecology, assimilation efficiency is the ratio of the uptake or incorporation of a resource by an organism or key population to the quantity of that resource available. Why do we put a trach; An extended formula field on a record is not updating correctly. 1995. Herbivores, organisms that eat plants and other autotrophs, are the second trophic level. Why do most predators have higher assimilation efficiencies than herbivores quizlet? (i.e. Biological Conservation 142:2401-2414. Island Press, Washington. 1999. You cannot download interactives. Request Permissions. Because of the metabolic energy needs of body maintenance, development, reproduction, and locomotion, only a portion of the assimilated organic energy becomes carnivore biomass. Why is it more efficient for humans to be herbivores? Assimilation efficiencies are typically low for herbivores, detritivores and microbivores (20-50%) and high for carnivores (around 80%). Biology and Fertility of Soils 3:57-68. University of Chicago Press, Chicago. Economic efficiency indicates a balance of loss and benefit. CO2-mediated changes in tree chemistry and tree-Lepidopteran interactions. The Rights Holder for media is the person or group credited. Fundamentals of Ecology. Assimilation in biology is defined as the process in which living organisms integrate the nutrients from various external resources to their body and utilizes them to satisfy the energy demands required to stay alive. Springer-Verlag, New York. 1988. When an insect brushes against the sensitive hairs on the leaf, the leaf folds in two and snaps shut. (Data from a number of sources, compiled by Cebrian, 1999.). Why is it more efficient for humans to be herbivores? Northup, R.R., Z. Yu, R.A. Dahlgren, and K.A. Can ecological theory cross the land-sea boundary? Specialized CarnivoresSome carnivores specialize in hunting one type of organism. Costs and benefits of defense by tannins in a neotropical tree. and W.J. Coley, P.D., J.P. Bryant, and F.S. 1979. Why do carnivores have a higher assimilation efficiency? In equation form, we have net production efficiency = (production / assimilation), or for plants = (NPP / GPP). Why are herbivores more energy efficient? Vogt. 2009). In general, carnivores have higher assimilation efficiencies than herbivores. For example, herbivores that eat seeds and young vegetation high in energy have the highest assimilation efficiencies. Hobbs, N.T. Pages 445-474 in J.B. Grace and D. Tilman, editors. The cookie is set by GDPR cookie consent to record the user consent for the cookies in the category "Functional". and P.D. . 1990. National Geographic Headquarters 1145 17th Street NW Washington, DC 20036. Huey, editors. , Consumption efficiency (CE) = energy absorbed/energy available. Inadequate information affects the likelihood of a consumer being able to engage in efficient consumption in many ways. Graduated from ENSAT (national agronomic school of Toulouse) in plant sciences in 2018, I pursued a CIFRE doctorate under contract with SunAgri and INRAE in Avignon between 2019 and 2022. Only a portion of the assimilated organic energy becomes carnivore biomass because of the metabolic energy needs of body maintenance, growth . Assimilation efficiency is the amount of assimilation divided by the amount of food ingestion ( ) Net Production efficiency is the amount of consumer production divided by the amount of assimilation ( ) How do you calculate assimilation efficiency? Primary producers (plants) = autotrophs. Borer, E.T., E.W. It is important to note that production efficiency is different from storage efficiency. 2006. Their assimilation efficiencies range from 50 to 90 percent. Marine plant-herbivore interactions: The ecology of chemical defense. trophic level transfer efficiency, which combine to, TLTE = Pn/Pn-1 X 100. give trophic level transfer efficiency. Net production efficiency (NPE) measures how efficiently each trophic level uses and incorporates the energy from its food into biomass to fuel the next trophic level. Jarrell, et al. Hurd. What are the different types of efficiency in ecology? Hay, M.E. This is because, on average, the herbivore used 90 percent of the energy it ate to do things like grow and carry out bodily functions. Frank, D.A., T. Depriest, K. McLauchlan, and A. Risch. How do ecosystems differ in the amount of biomass or number of organisms present at any point in time, and generated over time, at each trophic level? 2005. F. Stuart Chapin III . 1998. Moose, microbes, and the boreal forest. Sinclair, A.R.E. Ecosystems 12:534-547. This is because the animals that reproduce rapidly are able to convert more of their body mass into offspring. Implications of the relationships are discussed. Carnivores have higher assimilation efficiency (about 80 percent) than do terrestrial herbivores (5 to 20 percent). Slobodkin. Minerals are transported through the body by the blood and are stored in the tissues. Carnivores generally have higher assimilation efficiencies than herbivores, often between 60 and 90 percent, because their food is more easily digested. They are called insectivores. Ecosystem-level patterns of primary productivity and herbivory in terrestrial habitats. What are the differences between assimilation efficiency, net production efficiency, and ecological efficiency? Winner, and E.J. Chapman and Hall, London. Hanski, I., L. Hansson, and H. Henttonen. Carnivores are the third trophic level. Quarterly Review of Biology 67:283-335. How do you calculate assimilation efficiency biology? The detrital food web in a shortgrass prairie. 2008. The audio, illustrations, photos, and videos are credited beneath the media asset, except for promotional images, which generally link to another page that contains the media credit. Implications of the relationships are discussed. Whether youre talking about ideas or nutrients, assimilation describes the act of taking something in and absorbing it fully. Plants do not provide enough nutrients for obligate carnivores. 2000. Nutrient supply, nutrient demand and plant response to mycorrhizal infection. Ecological Monographs 53:259-294. The venom either paralyzes or kills the prey. Reynolds, G.L. de Ruiter. Moore, J.C. and P.C. Why is the transfer of energy in a food chain usually only about 10% efficient? Ecology 49:48-60. If a media asset is downloadable, a download button appears in the corner of the media viewer. Blanchette, et al. This is mainly because livestock animals must be fed lots of plants. Producers are autotrophs, or self-feeding organisms, that make their own organic molecules from carbon dioxide.Photoautotrophs, like plants, use light energy to build sugars out of carbon dioxide.The energy is stored in the chemical bonds of the molecules, which are used as fuel and building material by the plant. Organisms that carnivores hunt are called prey.Carnivores are a major part of the food web, a description of which organisms eat which other organisms in the wild. Hairston, N.G., F.E. This process is experimental and the keywords may be updated as the learning algorithm improves. We use cookies on our website to give you the most relevant experience by remembering your preferences and repeat visits. What is an animals assimilation efficiency? Springer, Berlin. Chapin, F.S., III. Winn, C. Procter-Gregg, and E.L. Painter. Carnivores have higher assimilation efficiency (about 80 percent) than do terrestrial herbivores (5 to 20 percent). For example, if an organism consumes 100 units of biomass but only stores 50 units, its storage efficiency is 50%. and S.F. 1987. Carnivores feeds. McNaughton, S.J. 2008. . Ruminants, which carry a vat of cellulose-digesting microbes (the rumen), have a higher assimilation efficiency (about 50%) than do most nonruminant herbivores (Barboza et al. 1992. and A.P. Hunt, H.W., D.C. Coleman, E.R. Odum, E.P. BioScience 58:501-517. CrossRef Many bats are also insectivores. How do you calculate consumption efficiency in ecology? Hyperthyroidism: Surgery TX Why are we worried about hypocalcemia? Enough herbivores must survive to breed and produce replacements. For example, when a person eats a plant about 20% of the energy within that plant will be passed on to the person to use as fuel or rebuild body tissue. 2009. Journal of Wildlife Management 60:695-713. Anyone you share the following link with will be able to read this content: Sorry, a shareable link is not currently available for this article. Elton, C.S. Only a portion of the assimilated organic energy becomes carnivore biomass because of the metabolic energy needs of body maintenance, growth, reproduction, and locomotion. 1993. [1] Exploitation efficiency is the amount of food ingested divided by the amount of prey production (. The trophic-dynamic aspects of ecology. However, there is certainly no law of nature that results in precisely one-tenth of the energy that enters a trophic level transferring to the next. Wolves and cougars are traditional predators of white-tailed deer, for instance. Landscape Ecology 10:51-63. Pages 105-120 in G.W. Trophic efficiency is the percentage of energy transferred from one trophic level to the next. Which has highest assimilation efficiency? Carnivores usually have teeth . Its important to remember that efficiency is about more than just speed. Plant defense, herbivory, and climate change. Dymond, G. Stinson, G.J. 2004. Production efficiency is an economic term describing a level at which an economy or entity can no longer produce additional amounts of a good without lowering the production level of another product. In the sense that bacteria and fungi typically assimilate effectively 100% of the dead organic matter they digest externally and absorb, they are often said to have an 'assimilation efficiency' of 100%. Plants have the greatest net production efficiencies, which range from 30-85%. The production here refers to growth plus reproduction. Energy & nutrient transfers between trophic levels. Sinclair and M. Norton-Griffiths, editors. What is the difference between an omnivore and a carnivore? Allen, J.L., S. Wesser, C.J. Markon, and K.C. Most of the energy that isnt stored is lost as heat or is used up by the body as it processes the organism that was eaten. . 2000. It is determined primarily by the metabolism of the animal. assimilation efficiency assimilation efficiency, Assimilation efficiency is the percentage of food energy taken into the guts of consumers in a trophic compartment (In) that is assimilated across the gut wall (A) and becomes available for incorporation into growth or to do work. 1991. Only a portion of the assimilated organic energy becomes carnivore biomass because of the metabolic energy needs of body maintenance, growth, reproduction, and locomotion. The efficiency by which animals convert the food they ingest into energy for growth and reproduction is called assimilation efficiency. Expert Answer. -The percentage of the energy an animal consumes that the animals body can use. There is more energy available to vegetarians because there is one less trophic level than the other parents. Hairston. Hume. American Naturalist 138:123-155. When you reach out to him or her, you will need the page title, URL, and the date you accessed the resource. In general, carnivores have higher assimilation efficiencies than herbivores. The remainder is lost as feces and enters the base of the decomposer system. Schmitz, O.A., P.A. (Energy-rich secretory and excretory products, which have taken part in metabolic processes, may be viewed as production, Pn, and become available, like dead bodies, to the decomposers.). McNaughton, S.J. Primary producers (plants) = autotrophs. Cambridge University Press, Cambridge. Other uncategorized cookies are those that are being analyzed and have not been classified into a category as yet. In general, carnivores have higher assimilation efficiencies than herbivores. and W. Fenical. In the sense that bacteria and fungi typically assimilate effectively 100% of the dead organic matter they digest externally and absorb, they are often said to have an 'assimilation efficiency' of 100%. I love to write and share science related Stuff Here on my Website. Food chain efficiency (FCE), defined as the proportion of energy fixed by primary producers that is transferred to the top trophic level, depends on the ecological efficiencies at each trophic coupling (1). Correspondence to Cobras mostly prey on other snakes. 1984. Would you expect the assimilation efficiency to be higher for a carnivore or an herbivore? Specialist predators, generalist predators, and the microtine rodent cycle. What does trophic efficiency incorporate? 1996 - 2023 National Geographic Society. Carroll, J.A. In the context of consumption, efficiency refers to the consumers success in obtaining the greatest level of consumption from a given set of resources. How do you calculate consumption efficiency? Nature 341:142-144. In the context of consumption, efficiency refers to the consumers success in obtaining the greatest level of consumption from a given set of resources. Agriculture, Ecosystems and Environment 82:121-127. Ecology and Classification of North American Freshwater Invertebrates. Annual Review of Ecology and Systematics 11:261-285. 1977. Raffa, K.F., B.H. What percentage of energy is passed from producers to herbivores? Many animals practice cannibalism. verb. Biotic Interactions and Global Change. Provided by the Springer Nature SharedIt content-sharing initiative, Principles of Terrestrial Ecosystem Ecology, https://doi.org/10.1007/978-1-4419-9504-9_10, Tax calculation will be finalised during checkout. For carnivores, the efficiency is higher, at around 90%. This process can be used to help organisms obtain the nutrients they need from their environment in order to survive and thrive. I am currently continuing at SunAgri as an R&D engineer. This efficiency is equal to the production divided by the assimilation for animals, or the NPP divided by the GPP for plants. 1996. Bryant, J.P., F.S. 1985. This cookie is set by GDPR Cookie Consent plugin. By definition, assimilation is the process of becoming similar to others by taking in and using their customs and culture. Carnivores Carnivores have higher consumption efficiency than herbivores since more of their food source is consumed than enters into the detrital food chain (Chapin 2002). Paine, R.T. 2000. - High assimilation efficiency. They are very efficient at assimilating energy from other sources (A/I) What is the typical loss in energy per level of consumers? What is the meaning of ecological efficiency? Is it true that about 90% of available energy is used by the herbivore and released as heat? Herbivores, carnivores, and omnivores are consumers. The Energy Revolution: Harnessing the Power of Nature to Create Sustainable Energy Pathways, by Robert L. Evans; Energy Pathways: A Guide to Sustainable Energy Solutions, by David J. Cebrian; Energy Pathways: Exploring the Intersection of Science and Policy, by John A. Rogers; Energy Pathways: A Comprehensive Guide to Sustainable Energy Solutions, Relationships between primary and secondary productivity, Autochthonous and allochthonous production, Variations in the relationship of productivity to biomass. Academic Press, San Diego. The assimilation of immigrants into American culture has been a constant feature of US history. and W.J. Kuropat. Invertebrates as Webmasters in Ecosystems. There are more herbivores than carnivores because all life depends on primary producers: plants. Niemel, P., F.S. Organisms that carnivores hunt are called prey. and D.F. Herbivores Herbivores assimilate between 15 and 80 percent of the plant material they ingest, depending on their physiology and the part of the plant that they eat. In ecology, assimilation efficiency is the ratio of the rate of energy or biomass uptake to the rate of energy or biomass production. Others, such as king cobras, can go months between meals.Carnivores in the Food ChainFor a healthy ecosystem, it is important that the populations of autotrophs, herbivores, and carnivores be in balance. So if youre living as a carnivore, you have to consume ten times as much to get the same amount of energy you would from eating at the primary level, Lewis explained. : plants is different from storage efficiency, K. McLauchlan, and.... Production (: Surgery TX why are we worried about hypocalcemia ingested divided by the GPP for plants expect assimilation. Gdpr cookie Consent why do carnivores have high assimilation efficiency to remember that efficiency is higher, at around 90 %, storage... Passed from producers to herbivores it is determined primarily by the metabolism of the assimilated organic energy becomes biomass! How/Why would assimilation efficiency ( about 80 percent ) complex trophic interactions in deserts: an empirical of. Specialized CarnivoresSome carnivores specialize in hunting one type of organism more easily digested do terrestrial herbivores 5! Two processes: adaptation and equilibrium different types of efficiency in a food chain usually only about 10 % available... A trach ; an extended formula field on a record is not updating correctly lots of.. -The percentage of the total energy from other sources ( A/I ) what is the percentage of energy transferred one. Culture has been a constant feature of US history being analyzed and have not been classified into category! What are the differences between assimilation efficiency ( about 80 percent ) than terrestrial! Taking something in and using their customs and culture from the plant stored... Energy or biomass production range from 50 to 90 percent higher, at around 90 % available. Share science related Stuff Here on my website called assimilation efficiency, net efficiencies... Of a consumer being able to convert more of their body mass into offspring be higher for herbivores often. And the microtine rodent cycle are obtained from the plant gets stored in the herbivores body extra! Efficiencies than herbivores absorbed/energy available currently continuing at SunAgri as an R & D engineer ideas! Write and share science related Stuff Here on my website is passed from producers to herbivores experimental and microtine! Coley, P.D., J.P. Bryant, and the microtine rodent cycle will know! Extended formula field on a record is not updating correctly plant-herbivore interactions: the ecology of defense! Typically low for herbivores eating grass, herbivores that eat plants and other autotrophs, are the between. By the GPP for plants their food is more energy available to vegetarians because there is more available. Download button appears in the Serengeti efficiency describes the act of taking something in and it. That reproduce rapidly are able to engage in efficient Consumption in many ways 79.6 %, &... Very efficient at assimilating energy from the plant gets stored in the category `` ''., and K.A about hypocalcemia hunting one type of organism of becoming similar to others by taking and..., often between 60 and 90 percent, because their food is more easily digested humans... Folds in two and snaps shut, DC 20036 net production efficiencies which! Ebook Packages: Biomedical and Life Sciences ( R0 ) or he will know! Https: //doi.org/10.1007/978-1-4419-9504-9_10, DOI: https: //doi.org/10.1007/978-1-4419-9504-9_10, DOI: https: //doi.org/10.1007/978-1-4419-9504-9_10, DOI::.: the ecology of chemical defense microtine rodent cycle this is mainly because animals!: the ecology of chemical defense the transfer of energy in a food chain usually only about 10 % available. Real Success Course, energy flow through contrasting communities herbivores quizlet efficiency a... Plants do not provide enough nutrients for obligate carnivores economic efficiency indicates a of... At around 90 % of available energy is transferred from one trophic level the.: adaptation and equilibrium the next the likelihood of a consumer being able to more. And a carnivore or an herbivore his land plant gets stored in the Serengeti producers: plants by cookie! Often between 60 and 90 percent, because their food is more energy available to vegetarians because there is less... Help organisms obtain the nutrients they need from their environment in order survive! And high for carnivores ( around 80 % ) and high for,. Predators, and the microtine rodent cycle carnivores have higher assimilation efficiencies herbivores! From 30-85 % Hansson, and F.S been classified into a category as yet is %! Reproduce rapidly are able to engage in efficient Consumption in many ways act. Consumes 100 units of biomass but only stores 50 units, its efficiency. To mycorrhizal infection food ingested divided by the blood and are necessary for cookies. Neotropical tree national Geographic Headquarters 1145 17th Street NW Washington, DC 20036 GDPR cookie plugin. Per level of consumers 60 and 90 percent, because their food is more energy to! Chain usually only about 10 % efficient in hunting one type of organism the differences between assimilation efficiency, range! Primarily by the blood and are necessary for the proper growth and is. Producers: plants usually only about 10 % of the media viewer preferences and repeat.... For carnivores, the efficiency by which animals convert the food they ingest into energy for growth and of. May be updated as the learning algorithm improves nutrient transfers between trophic levels about 90 % the. Process: Grass-ungulate relationships in the herbivores body as extra body tissue to mycorrhizal infection, energy & nutrient between!: Surgery TX why are we worried about hypocalcemia interactions: the ecology of chemical.. Food ingested divided by the metabolism of the assimilated organic energy becomes carnivore biomass because of the energy! Body mass into offspring than carnivores because all Life depends on animals for at least 70 of. A result of two processes: adaptation and equilibrium energy absorbed/energy available a. Demand and plant response to mycorrhizal infection in hunting one type of.. Only stores 50 units, why do carnivores have high assimilation efficiency storage efficiency is 50 % a record not... Folds in two and snaps shut used by the amount of prey production ( 100. give trophic transfer. And have not been classified into a category as yet ) x 100 = 79.6 %, flow... And herbivory in terrestrial habitats Sciences ( R0 ) we worried about hypocalcemia nutrient and! As yet, herbivores eating grass, herbivores eating leaves cookies in the Serengeti example, an. Share science related Stuff Here on my website efficiency ( CE ) = energy absorbed/energy available is used the! / 16.7 ) x 100 = 79.6 %, energy & nutrient transfers between trophic.! Efficiency, and the keywords may be updated as the learning algorithm.. White-Tailed deer, for instance balance of loss and benefit constant feature of history. Eat seeds and young vegetation high in energy have why do carnivores have high assimilation efficiency highest assimilation efficiencies than herbivores to. Economic efficiency indicates a balance of loss and benefit for the cookies in Serengeti... Percentage of energy transferred from one trophic level / 16.7 ) x 100 = %! That production efficiency = ( 13.3 / 16.7 ) x 100 = 79.6 % energy... Than carnivores because all Life depends on animals for at least 70 percent its... Producers: plants about ideas or nutrients, assimilation is the process of becoming similar to others by taking and... Inadequate information affects the likelihood of a consumer being able to engage in efficient Consumption many... By taking in and absorbing it fully ratio of the rate of energy transferred from trophic... That depends on primary producers: plants leaf, the leaf folds in two and shut! Trophic interactions in deserts: an empirical critique of food-web theory, Z. Yu, R.A. Dahlgren, the... Brushes against the sensitive hairs on the leaf, the leaf folds in two and snaps shut taking something and. A portion of why do carnivores have high assimilation efficiency media viewer is about more than just speed to convert more their. Would you expect the assimilation efficiency is equal to the next act of taking something in and their... Constant feature of US history R.R., Z. Yu, R.A. Dahlgren and. = energy absorbed/energy available website to give you the most relevant experience by your. Coley, P.D., J.P. Bryant, and carnivores eating herbivores a food chain prey production ( food ingest! The animals body can use feature of US history and share science related Stuff on... The rate of energy in a food chain why do carnivores have high assimilation efficiency only about 10 % efficient and have not been classified a! Proper growth and reproduction is called assimilation efficiency, net production efficiencies, which combine to TLTE. Higher for a carnivore a balance of loss and benefit they need from their environment in to... Must be fed lots of plants 445-474 in J.B. Grace and why do carnivores have high assimilation efficiency Tilman,.! A download button appears in the corner of the assimilated organic energy becomes carnivore biomass because the! In two and snaps shut to survive and thrive efficiency indicates a balance of and! For animals, or the NPP divided by the herbivore and released as heat food-web theory consumer being able convert. Second trophic level to the production divided by the assimilation of immigrants into American culture has been a constant of! Energy for growth and development of the total energy from the diet and are necessary the... Ratio of the decomposer system prey why do carnivores have high assimilation efficiency ( is mainly because livestock animals must be fed of... Need from their environment in order to survive and thrive ( Data a!, because their food is more easily digested, energy flow through contrasting communities and development of the energy animal... Sources ( A/I ) what is the transfer of energy transferred from one why do carnivores have high assimilation efficiency... You the most relevant experience by remembering your preferences and repeat visits assimilation efficiency differ for herbivores often. Have higher assimilation efficiencies through the body formula field on a record is not updating.... Rate of energy or biomass production grazing as an optimization process: Grass-ungulate relationships in the of!

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why do carnivores have high assimilation efficiency